Nationality
Dutch

Year of Birth
1957

• M.Sc. in Biology, University of Utrecht, NL

• M.D. University of Utrecht, NL

• Ph.D. University of Utrecht, NL

1985-1989
Postdoctoral Fellow Pathology, Cox Terhorst Lab at the Dana-Farber Cancer Institute, Harvard Medical School, Boston MA, USA

1989-1991
Assistant Professor, Department of Clinical Immunology, University of Utrecht, NL
Lecturer in Clinical Immunology

1991-2002
Professor and Chairman, Dept. of Immunology, Faculty of Medicine, University of Utrecht, NL

2002-2012
Head Director of the Hubrecht Institute, Royal Netherlands Academy of Arts and Sciences, Utrecht, NL

2002-present
Professor in Molecular Genetics, University of Utrecht, NL

2002-2022
Principal Investigator of a research group of ~25 scientists at the Hubrecht Institute, Utrecht, NL (continued as advisor beyond March 2022)

2012-2015
President of the Royal Netherlands Academy of Sciences, Utrecht, NL

2015-2019
Director Research of the Princess Máxima Center for Pediatric Oncology, Utrecht, NL

2015-2022
Principal lnvestigator of a research group at the Princess Máxima Center for Pediatric Oncology, Utrecht, NL (continued as advisor beyond 2022)

2005-2015
Member of the Scientific Advisory Board of Swiss Institute for Experimental Cancer Research (ISREC), Lausanne, CH

2006-2008
President of the International Society of Differentiation (ISD)

2012-2015
Member of the board of the American Association for Cancer Research

2014-2020
Chief Scientific Officer der HUB Organoids Technology, Utrecht, NL

2015-2021
Member of the Scientific Advisory Board of the Institute of Molecular Pathology (IMP), Vienna, AU

2016-present
Member of the Scientific Advisory Board of the Francis Crick Institute, London, UK

2016-2021
Member of the Scientific Advisory Board of Decibel Therapeutics, Boston, USA

2016-2022
Member of the Scientific Advisory Board and co-founder of Surrozen, San Francisco, USA

2016-2020
Member of the Scientific Advisory Board of Kallyope, New York, USA

2017-2018
President of the International Society for Stem Cell Research (ISSCR)

2018-2021
Member of the Scientific Advisory Board of Merus, Utrecht, NL

2019-2022
Member of the Board of Directors of Roche Holding Ltd.

2020-2022
Member of the Scientific Advisory Board and co-founder of Xilis, North Carolina, USA

2021-2022
Member of the Scientific Advisory Board and co co-founder of D1 Med, Shanghai, China

2021-2021
Member of the Scientific Advisory Board of Volastra, New York, USA

2000
Royal Netherlands Academy of Sciences (KNAW), Amsterdam, NL

2014
National Academy of Sciences of the USA, Washington, USA

2016
Academie des Sciences de I 'Institut de France, Paris, F

2017
Orden Pour le Mérite für Wissenschaften und Künste, Berlin, DE

2019
The Royal Society, London, UK

2019
The Royal Society, Edinburgh, UK